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- Professor Emeritus
- Department of Anatomy, Physiology, and Cell Biology, School of Veterinary Medicine
- Professor
- Department of Population Health and Reproduction, School of Veterinary Medicine
- Professor Emeritus
- Department of Psychology, College of Letters and Science
- Professor
- Department of Evolution and Ecology, College of Biological Sciences
- Professor
- Department of Neurobiology, Physiology and Behavior, College of Biological Sciences
